Photosynthetic organisms continually encounter fluctuations in their environment, particularly in the quantity of incident light. In this work we investigate especially acclimation changes concerning EET efficiency from LHCII to RC of PSII. Our results have suggested that acclimation changes of EET efficiency from LHCII to RC of PSII comes from macro-organization of PSII supercomplexes, which is larger for both plants grown under low light conditions. It has been demonstrated by bigger psi-type CD bands in the CD spectra of thylakoid membranes. We assume this, because together with a disturbance of macrodomains changes of EET efficiency from Chl b and Car to emitted Chl a have been lost.